When the evil mayor of Oakton decides to bulldoze Liberty Park and build a dangerous amusement park in its place, Surly Squirrel and his ragtag group of animal friends need to band together to save their home, defeat the mayor, and take back the park.

Surly, a curmudgeon, independent squirrel is banished from his park and forced to survive in the city. Lucky for him, he stumbles on the one thing that may be able to save his life, and the rest of park community, as they gear up for winter - Maury's Nut Store.